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A huge meteorite has plunged into the Great Valley, cutting off the water supply and causing tension among the once-peaceful dinosaurs. While the adults argue and turn against one another, Littlefoot and his friends venture out to find a new source of water, only to be constantly bullied by a trio of older dinosaurs. The journey becomes a fight for survival when the group is cornered by a pack of Velociraptors and trapped by a sudden wildfire.
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A plague of locusts strikes the Great Valley, decimating the plants and forcing the dinosaurs to migrate in search of food. When the adults in the herd begin fighting over the scarce resources, young Littlefoot and his friends Petrie, Cera, Ducky and Spike split off from the main group, becoming stranded on an island after their path was destroyed by a tsunami. As they search for a way off the island, they encounter an old acquaintance who may become friend or foe.
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Littlefoot and his friends find life in the Great Valley a bit too restrictive under the watchful eyes of their overprotective parents. Wishing to not be treated like babies anymore, the group chase after a pair of dangerous dinosaurs stealing an egg from Ducky's nest. When the egg they bring back hatches in the valley, the friends are surprised to find a baby Sharptooth. Rather than abandoning him, they name him Chomper and attempt to raise him as one of their own.
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